Chalivendra C. Subbaiah, Douglas S. Bush, Martin M. Sachs, Elevation of Cytosolic Calcium Precedes Anoxic Gene Expression in Maize Suspension-Cultured Cells, The Plant Cell, Vol. 6, No. 12 (Dec., 1994), pp. 1747-1762
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Subbaiah et al. (1994) studied this question.