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September 8, 2026Neurobiology of DiseaseOpen Access

Activity-dependent DNA methylation and demethylation: epigenetic regulators of learning and memory

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Authors

ZLZhehao LiZXZiyue XuXLXuefeng Li

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Overview

Review highlights activity-dependent DNA methylation dynamics in the brain, suggesting novel epigenetic targets for treating cognitive decline and Alzheimer's disease.

Key Points

  • This review synthesizes how activity-dependent DNA methylation and demethylation orchestrate persistent transcriptional programs required for synaptic plasticity, learning, and memory.
  • Synthesized molecular evidence on canonical cytosine modifications (5mC, 5hmC, 5fC, 5caC) and their modifying enzymes, including DNMTs, TETs, and TDG.
  • Evaluated emerging literature on non-canonical modifications (6mA and 4mC) alongside recent advances in high-resolution brain epigenomic mapping technologies.
  • Dynamic oxidative turnover of 5-methylcytosine serves as a primary driver of activity-regulated transcription essential for neuronal plasticity and memory consolidation.
  • Dysregulation of DNA methylation and demethylation networks is directly linked to transcriptional dysfunction and progressive cognitive decline in Alzheimer's disease.
  • Emerging high-resolution mapping technologies enable locus- and cell-type-specific tracking of epigenetic marks, providing tools to discover targeted cognitive therapeutics.

Cite This Study

Li et al. (2026) studied this question.

synapsesocial.com/papers/6a9fd72a58e84d0ff5b45af9https://doi.org/10.1016/j.nbd.2026.107596
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