E. John Ainsworth, Lola S. Kelly, Lynn J. Mahlmann, John C. Schooley, R. H. Thomas, Jerry Howard, Edward L. Alpen, Response of Colony-Forming Units-Spleen to Heavy Charged Particles, Radiation Research, Vol. 96, No. 1 (Oct., 1983), pp. 180-197
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Ainsworth et al. (1983) studied this question.