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Alvan A. Karlin, Sheldon I. Guttman, Stephen L. Rathbun, Spatial Autocorrelation Analysis of Heterozygosity and Geographic Distribution in Populations of Desmognathus fuscus (Amphibia: Plethodontidae), Copeia, Vol. 1984, No. 2 (May 1, 1984), pp. 343-356
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Karlin et al. (1984) studied this question.
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