John C. Larkin, David G. Oppenheimer, Alan M. Lloyd, Ellen T. Paparozzi, M. David Marks, Roles of the GLABROUS1 and TRANSPARENT TESTA GLABRA Genes in Arabidopsis Trichome Development, The Plant Cell, Vol. 6, No. 8 (Aug., 1994), pp. 1065-1076
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Larkin et al. (1994) studied this question.
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