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May 14, 2009Proceedings of the National Academy of Sciences156 citationsOpen Access

The small G-proteins Rac1 and Cdc42 are essential for myoblast fusion in the mouse

EVElena VasyutinaBMBenedetta MartarelliCBCord Brakebusch

Key Result

Conditional mutagenesis of Rac1 or Cdc42 in mice significantly impaired myoblast fusion, reducing the fusion index from 52% in controls to 8% and 26%, respectively.

Structured PICO

P
Population
Mouse embryos with conditional mutations in Rac1 or Cdc42 used to study myoblast fusion during development.
I
Intervention
Conditional mutagenesis of Rac1 and Cdc42
C
Comparator
Control embryos (Rac1flox/+;Lbx1cre or Cdc42flox/+;Lbx1cre)
O
Outcome
Myoblast fusion in vivo and in vitro (fusion index and recruitment of cytoskeletal proteins to contact sites)surrogate

Rac1 and Cdc42 are essential for myoblast fusion in mice, functioning in a nonredundant manner to recruit vinculin and cytoskeletal proteins to contact sites.

Main Result

Absolute Event Rate: 8% vs 52%

Limitations

  • Animal model findings may not fully translate to human biology
  • The Cdc42 phenotype might reflect a greatly reduced but not completely absent Rac1 activity

Abstract

Rac1 and Cdc42 are small G-proteins that regulate actin dynamics and affect plasma membrane protrusion and vesicle traffic. We used conditional mutagenesis in mice to demonstrate that Rac1 and Cdc42 are essential for myoblast fusion in vivo and in vitro. The deficit in fusion of Rac1 or Cdc42 mutant myoblasts correlates with a deficit in the recruitment of actin fibers and vinculin to myoblast contact sites. Comparison of the changes observed in mutant myogenic cells indicates that Rac1 and Cdc42 function in a nonredundant and not completely overlapping manner during the fusion process. Our genetic analysis demonstrates thus that the function of Rac in myoblast fusion is evolutionarily conserved from insects to mammals and that Cdc42, a molecule hitherto not implicated in myoblast fusion, is essential for the fusion of murine myoblasts.

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Cite This Study

Vasyutina et al. (2009) studied Muscle development. Conditional mutagenesis of Rac1 and Cdc42 vs. Control mice (Rac1flox/+; Lbx1cre or Cdc42flox/+; Lbx1cre) was evaluated on Fusion index (percentage of myogenic nuclei present in multinucleated cells) at E14.5. Conditional mutagenesis of Rac1 or Cdc42 in mice significantly impaired myoblast fusion, reducing the fusion index from 52% in controls to 8% and 26%, respectively.

synapsesocial.com/papers/6aa15fb11e707e103972dfbfhttps://doi.org/10.1073/pnas.0902501106
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