Anton A. Sanderfoot, Sondra G. Lazarowitz, Cooperation in Viral Movement: The Geminivirus BL1 Movement Protein Interacts with BR1 and Redirects It from the Nucleus to the Cell Periphery, The Plant Cell, Vol. 7, No. 8 (Aug., 1995), pp. 1185-1194
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Sanderfoot et al. (1995) studied this question.