Yong-Qiang Gu, Caimei Yang, Venkatappa K. Thara, Jianmin Zhou, Gregory B. Martin, Pti4 Is Induced by Ethylene and Salicylic Acid, and Its Product Is Phosphorylated by the Pto Kinase, The Plant Cell, Vol. 12, No. 5 (May, 2000), pp. 771-785
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Gu et al. (2000) studied this question.