Jane E. Dorweiler, Charles C. Carey, Kenneth M. Kubo, Jay B. Hollick, Jerry L. Kermicle, Vicki L. Chandler, mediator of paramutation1 Is Required for Establishment and Maintenance of Paramutation at Multiple Maize Loci, The Plant Cell, Vol. 12, No. 11 (Nov., 2000), pp. 2101-2118
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Dorweiler et al. (2000) studied this question.