Gilles Houle, Spatiotemporal Patterns in the Components of Regeneration of Four Sympatric Tree Species -- Acer Rubrum, A. Saccharum, Betula Alleghaniensis and Fagus Grandifolia, Journal of Ecology, Vol. 82, No. 1 (Mar., 1994), pp. 39-53
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Gilles Houle (1994) studied this question.