The distribution of phytoplankton was studied in Lake Lanao, Philippines. In calm weather, the species segregate vertically with motile species and buoyancy‐regulating species above other species in the water column. On a distance scale of 12.5 km, horizontal variability yields a coefficient of variation of 23.6% for total biomass, 30.2% for net primary production, and an average of 30.3% for individual species biomass after removal of error variances. Variabilities on a scale of 1 km are about half as large. Horizontal patchiness of individual species is not related to their abundance, surface to volume ratio, morphotype, or taxon, but is negatively related to their size.
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William M. Lewis (1978) studied this question.
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