D. A. Wardle, G. W. Yeates, R. N. Watson, K. S. Nicholson, Development of the Decomposer Food-Web, Trophic Relationships, and Ecosystem Properties during a Three-Year Primary Succession in Sawdust, Oikos, Vol. 73, No. 2 (Jun., 1995), pp. 155-166
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Wardle et al. (1995) studied this question.
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