In accelerating the elongation of excised Avena coleoptile cylinders, the effect of oxalic acid was found to be additive to that of indole-3-acetic acid. Elastic and plastic extensions of the coleoptile cylinder were measured by using the plasmolytic method. Both indole-3-acetic acid and oxalic acid increased the elastic extension quickly, while their effect in increasing the plastic extension appeared 2 hours after their application. The mechanism of auxin action is discussed in connection with the effect of removal of calcium from the pectic substance of the primary cell wall.
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Yoshio Masuda (1961) studied this question.