Answers are grounded in this paper and cite their sources.
The shape of the cranium of the fifteen million year old (Astaracian, MN zone 6) Eotragus sansaniensis from Sansan most closely resembles that of Tetracerus quadricornis (Boselaphini) but also resembles that of Cephalophus natalensis (Cephalophini), while E. sansaniensis horn cores are most similar to those of Boselaphus tragocamelus (Boselaphini). Morphologically, E. sansaniensis is one of the most primitive bovids known and is placed in the Boselaphini.
No takes yet. Share an insight, caveat, or question.
Solounias et al. (1992) studied this question.
Synapse has enriched 10 related papers. Consider them for comparative context: