Nien-Chen Huang, Chien-Sung Chiang, Nigel M. Crawford, Yi-Fang Tsay, CHL1 Encodes a Component of the Low-Affinity Nitrate Uptake System in Arabidopsis and Shows Cell Type-Specific Expression in Roots, The Plant Cell, Vol. 8, No. 12 (Dec., 1996), pp. 2183-2191
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Huang et al. (1996) studied this question.