Leucine and isoleucine conjugates of 1-oxo-2,3-dihydro-indene-4-carboxylic acid (1-oxo-ICA) such as 2a-e are powerful elicitors of volatile biosynthesis in the Lima bean ( Phaseolus lunatus ). The compounds are structurally and, with a few exceptions, functionally related to coronatin 1, a phytotoxin from pathovars of Pseudomonas spp. Coronatin 1 and the conjugates of 1-oxo-ICA 2a-e seem to mimic signals from the octadecanoid signalling pathway, like, for example, amino acid conjugates of jasmonic acid.
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Thomas Krumm (1996) studied this question.