A scanning electron microscope study showed indentations by the follicle wall cells into the feather shaft. In other areas altered cells or keratinised cell fragments form bridges which join the feather to the follicle. 2. In the early stage of growth the feather shaft is held in place by cell indentations, whereas later, bridges are seen which probably hold the feather more firmly.
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ANGEL et al. (1982) studied this question.
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