The studies reported in this paper were made as opportunities and materials offered during the course of investigations of the leaf rust of the grasses, the results of which have been given elsewhere (1933). A number of the cultures were made possible through the kindness of the late Mr. E. Bethel, to whom the writer is specially indebted for material and clues concerning connections based on his discriminating field observations.
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E. B. Mains (1933) studied this question.
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