Bart Claes, Rudy Dekeyser, Raimundo Villarroel, Marc Van den Bulcke, Guy Bauw, Marc Van Montagu, Allan Caplan, Characterization of a Rice Gene Showing Organ-Specific Expression in Response to Salt Stress and Drought, The Plant Cell, Vol. 2, No. 1 (Jan., 1990), pp. 19-27
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Claes et al. (1990) studied this question.