S. J. Hall, M. R. Robertson, D. J. Basford, R. Fryer, Pit-Digging by the Crab Cancer pagurus: A Test for Long-Term, Large-Scale Effects on Infaunal Community Structure, Journal of Animal Ecology, Vol. 62, No. 1 (Jan., 1993), pp. 59-66
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Hall et al. (1993) studied this question.
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