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M. V. Andrews, D. D. Gilbertson, M. Kent, P. A. Mellars, Biometric Studies of Morphological Variation in the Intertidal Gastropod Nucella lapillus (L): Environmental and Palaeoeconomic Significance, Journal of Biogeography, Vol. 12, No. 1 (Jan., 1985), pp. 71-87
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Andrews et al. (1985) studied this question.
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