Guy Naulleau, Xavier Bonnet, Myriam Vacher-Vallas, Richard Shine, Oliver Lourdais, Does Less-than-Annual Production of Offspring by Female Vipers (Vipera aspis) Mean Less-than-Annual Mating?, Journal of Herpetology, Vol. 33, No. 4 (Dec., 1999), pp. 688-691
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Naulleau et al. (1999) studied this question.
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