Maureen Stanton, Helen J. Young, Norman C. Ellstrand, Janet M. Clegg, Consequences of Floral Variation for Male and Female Reproduction in Experimental Populations of Wild Radish, Raphanus sativus L., Evolution, Vol. 45, No. 2 (Mar., 1991), pp. 268-280
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Stanton et al. (1991) studied this question.