Steffen Reinbothe, Christiane Reinbothe, Nikolai Lebedev, Klaus Apel, PORA and PORB, Two Light-Dependent Protochlorophyllide-Reducing Enzymes of Angiosperm Chlorophyll Biosynthesis, The Plant Cell, Vol. 8, No. 5 (May, 1996), pp. 763-769
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Reinbothe et al. (1996) studied this question.