Alain F. Tissier, Sylvestre Marillonnet, Victor Klimyuk, Kanu Patel, Miguel Angel Torres, George Murphy, Jonathan D. G. Jones, Multiple Independent Defective Suppressor-mutator Transposon Insertions in Arabidopsis: A Tool for Functional Genomics, The Plant Cell, Vol. 11, No. 10 (Oct., 1999), pp. 1841-1852
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Tissier et al. (1999) studied this question.