J. J. Boomsma, G. A. Van Der Lee, T. M. Van Der Have, On the Production Ecology of Lasius niger (Hymenoptera: Formicidae) in Successive Coastal Dune Valleys, Journal of Animal Ecology, Vol. 51, No. 3 (Oct., 1982), pp. 975-991
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Boomsma et al. (1982) studied this question.
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