Charles B. Beard, Jerry F. Butler, Ellis C. Greiner, In vitro Growth Characterization and Host-Parasite Relationship of Leptomonas pulexsimulantis n. sp., a Trypanosomatid Flagellate of the Flea Pulex simulans, The Journal of Parasitology, Vol. 75, No. 5 (Oct., 1989), pp. 658-668
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Beard et al. (1989) studied this question.
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