J. M. Dick, P. G. Jarvis, R. R. B. Leakey, Influence of Male and Female Cones on Needle CO2 Exchange Rates of Field-Grown Pinus contorta Doug. Trees, Functional Ecology, Vol. 5, No. 3 (1991), pp. 422-432
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Dick et al. (1991) studied this question.
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