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Mauro Santos, Daniel J. Borash, Amitabh Joshi, Nira Bounlutay, Laurence D. Mueller, Density-Dependent Natural Selection in Drosophila: Evolution of Growth Rate and Body Size, Evolution, Vol. 51, No. 2 (Apr., 1997), pp. 420-432
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Santos et al. (1997) studied this question.
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