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C. J. Bult, J. A. Sweere, E. A. Zimmer, Cryptic Sequence Simplicity, Nucleotide Composition Bias, and Molecular Coevolution in the Large Subunit of Ribosomal DNA in Plants: Implications for Phylogenetic Analyses, Annals of the Missouri Botanical Garden, Vol. 82, No. 2 (1995), pp. 235-246
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Bult et al. (1995) studied this question.
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