Jukka Jokela, Curtis M. Lively, Jennifer A. Fox, Mark F. Dybdahl, Flat Reaction Norms and "Frozen" Phenotypic Variation in Clonal Snails (Potamopyrgus antipodarum), Evolution, Vol. 51, No. 4 (Aug., 1997), pp. 1120-1129
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Jokela et al. (1997) studied this question.