Robert M. Hughes, Eric Rexstad, Carl E. Bond, The Relationship of Aquatic Ecoregions, River Basins and Physiographic Provinces to the Ichthyogeographic Regions of Oregon, Copeia, Vol. 1987, No. 2 (May 13, 1987), pp. 423-432
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Hughes et al. (1987) studied this question.
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