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Orkney voles Microtus arvalis show avoidance of red fox Vulpes vulpes faecal odours, both in the laboratory and in the wild. Since these voles have been isolated from foxes since Neolithic times it is likely that these responses are innate rather than learned. When applied to young Scots pine saplings, the fox odour inhibited feeding by voles when alternative foods were available.
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Calder et al. (1991) studied this question.
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