Robert B. Giannattasio, David M. Spooner, A Reexamination of Species Boundaries Between Solanum megistacrolobum and S. toralapanum (Solanum sect. Petota, series Megistacroloba): Morphological Data, Systematic Botany, Vol. 19, No. 1 (Jan. - Mar., 1994), pp. 89-105
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Giannattasio et al. (1994) studied this question.
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