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F. N. Young, C. C. Goff, An Annotated List of the Arthropods Found in the Burrows of the Florida Gopher Tortoise, Gopherus polyphemus (Daudin), The Florida Entomologist, Vol. 22, No. 4 (Dec., 1939), pp. 53-62
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Young et al. (1939) studied this question.
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