Stewart B. Peck, New Montane Ptomaphagus Beetles from New Mexico and Zoogeography of Southwestern Caves (Coleoptera; Leiodidae; Catopinae), The Southwestern Naturalist, Vol. 23, No. 2 (Mar. 20, 1978), pp. 227-238
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Stewart B. Peck (1978) studied this question.
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