Authors
F. Mestres, G. Pegueroles, A. Prevosti, L. Serra, Colonization of America by Drosophila subobscura: Lethal Genes and the Problem of the O5 Inversion, Evolution, Vol. 44, No. 7 (Nov., 1990), pp. 1823-1836
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Mestres et al. (1990) studied this question.