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Tod F. Stuessy, Daniel J. Crawford, Clodomiro Marticorena, Patterns of Phylogeny in the Endemic Vascular Flora of the Juan Fernandez Islands, Chile, Systematic Botany, Vol. 15, No. 2 (Apr. - Jun., 1990), pp. 338-346
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Stuessy et al. (1990) studied this question.
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