Adolfo Cordero, Density-Dependent Mating Success and Colour Polymorphism in Females of the Damselfly Ischnura graellsii (Odonata: Coenagrionidae), Journal of Animal Ecology, Vol. 61, No. 3 (Oct., 1992), pp. 769-780
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Adolfo Cordero (1992) studied this question.
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