A. P. Mamolos, G. K. Elisseou, D. S. Veresoglou, Depth of Root Activity of Coexisting Grassland Species in Relation to N and P Additions, Measured Using Nonradioactive Tracers, Journal of Ecology, Vol. 83, No. 4 (Aug., 1995), pp. 643-652
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Mamolos et al. (1995) studied this question.
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