Charles H. Willey, Gabriel C. Godman, Gametogenesis, Fertilization and Cleavage in the Trematode, Zygocotyle lunata (Paramphistomidae), The Journal of Parasitology, Vol. 37, No. 3 (Jun., 1951), pp. 283-296
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Willey et al. (1951) studied this question.
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