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The F1 intergeneric hybrids of Psathyrostachys huashanica Keng with Roegneria ciliaris (Trin.) Nevski was made and cytologically studied. Chromatin transfer through conjugation tube or conjugation opening before, during and after meiosis was observed in these hybrids. Consequently, unusual nuclear behaviors frequently observed include coenocytism, cell size variation and abnomal chromosome or nuclei number, non-synchronous or multipolar division, and delayed chromatin condensation. In addition, cytomixis was observed in early stages of micro-sporogenesis. These events could lead to spontaneous chromosome number doubling and, in turn, the origin of alloautoploid species.
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Yen et al. (1993) studied this question.
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