L. P. Ronse Decraene, E. Smets, The Systematic Relationship between Begoniaceae and Papaveraceae: A Comparative Study of Their Floral Development, Bulletin du Jardin botanique National de Belgique / Bulletin van de Nationale Plantentuin van België, Vol. 60, No. 3/4 (Dec. 31, 1990), pp. 229-273
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Decraene et al. (1990) studied this question.