Loren H. Rieseberg, Douglas E. Soltis, Jeffrey D. Palmer, A Molecular Reexamination of Introgression between Helianthus annuus and H bolanderi (Compositae), Evolution, Vol. 42, No. 2 (Mar., 1988), pp. 227-238
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Rieseberg et al. (1988) studied this question.
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