We sequenced parts of three mitochondrial genes in 67 great spotted woodpeckers Dendrocopos major taken from 17 sites across Eurasia and Japan. Two groups, differing by ca. 3% sequence divergence, were determined to have had independent evolutionary histories: samples from the southeast (Sakhalin, Primor'e, Hokkaido), and the rest of Eurasia. The southeastern group corresponds to the subspecies D. m. japonicus , which will likely merit species status upon study of all subspecies; at present it should be considered an evolutionarily significant unit. From the United Kingdom to the Russian North Pacific, no phylogeographic divisions were found, which is likely a result of recent range expansion following retreat of permafrost after the last ice age.
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Zink et al. (2002) studied this question.
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