Jeffrey R. Johansen, Larry L. St. Clair, Bruce L. Webb, Glen T. Nebeker, Recovery Patterns of Cryptogamic Soil Crusts in Desert Rangelands Following Fire Disturbance, The Bryologist, Vol. 87, No. 3 (Autumn, 1984), pp. 238-243
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Johansen et al. (1984) studied this question.
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