may be carried in specialized cuticular structures, termed mycangia, that can take a variety of forms (Francke-Grosmann, 1967.pp.141-205.In S. M. Henry (ed.), Symbiosis Vol.2., Academic Press, New York).Several species of Dendroctonus bark beetles possess thoracic (e.g., D. brevicomis
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Six et al. (1996) studied this question.