L. D. Marshall, J. N. McNeil, Spermatophore Mass as an Estimate of Male Nutrient Investment: A Closer Look in Pseudaletia unipuncta (Haworth) (Lepidoptera: Noctuidae), Functional Ecology, Vol. 3, No. 5 (1989), pp. 605-612
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Marshall et al. (1989) studied this question.
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