During 1975, 12 km south of Boothbay Harbor, soluble iron concentrations in the first 20 m of water were three times greater in October and November than in August and September, while particulate iron concentrations only increased transiently before the fall bloom. Nutrient enrichment experiments and chlorophyll a : cytochrome f1 ratios indicated that low iron concentrations limited phytoplankton populations. In August 1976, 1–4 km from Monhegan Island, increased iron concentrations from land runoff preceded a dinoflagellate bloom.
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Hilary E. Glover (1978) studied this question.