Ellen Kalmbach, Stephen C. Ramsay, Helmut Wendeln, Peter H. Becker, A Study of Neotropic Cormorants in Central Chile: Possible Effects of El Niño, Waterbirds: The International Journal of Waterbird Biology, Vol. 24, No. 3 (Dec., 2001), pp. 345-351
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Kalmbach et al. (2001) studied this question.
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